Mack A Beacon

Joined: Aug 9, 2020 Last Active: Feb 18, 2024 iNaturalist

We were all rookies at identification, and we learn as we go from others with knowledge. I want you to know that it is my pleasure to click "AGREE" with you're ID, I love people trying there best to get it right. Also I do see your thank you's but do to me doing 30-200 ID's most nights, I just don't have time to respond, but know I see it, and apriciate your thank you's.

If I moved your sample from your [genus/species] pick back to genus it is because there is not enough evidence of species so it must return to genus or even back to family. I have great respect for those that try this ID process.

Attention to members information you should know; The only away to be 100% sure of an ID is if someone like myself handles the sample and microscopically identifies it to species and or synonym. I have been handling Ephemeroptera daily since 2010 and I am personally connected to all the world authorities to help me if need. Identification by photo is problematic for many reasons, therefor latitude must always be considered. If you need help, email me.


Currently I am writing a revision paper for the genus Stenacron that addresses the interpunctatum complex. The biosystematics paper is also underway for the genus Leucrocuta 2025. As it stands there are no true larva descriptions across the Leucrocuta genus.


Heptageniidae quick link.
https://www.inaturalist.org/observations/identify?taxon_id=48006&place_id=97394

Leucrocuta quick link.
https://www.inaturalist.org/observations/identify?place_id=97394&taxon_id=177453


My Feb 2021 paper; Taxonomic Dissection charts of the larvae Maccaffertium and Stenonema.
https://www.academia.edu/45291722/Maccaffertium_and_Stenonema_Larva_diagnostic_charts

My Jan 2021 paper Heptagenia pulla larva.
https://www.academia.edu/45185541/A_deep_review_of_H_pulla_larva_and_its_variation

My December 2020 paper on Maccaffertium vicarium complex;
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubbeaconm2020bp1.pdf

My book on Stenacron 2020
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubbeaconm2020p1.pdf?fbclid=IwAR1Nx-uAY8iV0mq4-DjLBkPJDTWqIv9tj_BsnNRiQ0ACqR0WoTq_zfy1KNY

Area of specialties HEPTAGENIIDAE;
Anepeorus, Cinygma, Cinygmula, Ecdyonurus, Epeorus, Heptagenia, Ironodes, Leucrocuta, Maccaffertium, Macdunnoa, Nixe, Raptoheptagenia cruentata, Rhithrogena, Spinadis simplex, Stenacron,Stenonema.

Leucrocuta petersi larva description.
https://www.ephemeroptera-galactica.com/pubs/pub_b/pubbednarika1980p51.pdf

Last collections for leucrocuta jewetti
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmeyerm2007p65.pdf

Mayfly centeral
https://www.entm.purdue.edu/mayfly/na-species-list.php

Luke Jacobus must read.
https://www.washingtonpost.com/magazine/2022/09/19/mayfly-decline/

Traver JR; Edmunds GF Jr. 1967. A revision of the genus Thraulodes
https://www.ephemeroptera-galactica.com/pubs/pub_t/pubtraverj1967p349.pdf

Diphetor hageni + larva.
file:///C:/Users/HP/Downloads/EPA-260-R-08-014TaxonomicAidsforMid-AtlanticBenthicMacroinvertebrates.pdf

Provonchsa 1990 Caeniidae revision
https://www.ephemeroptera-galactica.com/pubs/pub_p/pubprovonshaa1990p801.pdf

Provonchsa & MaCafferty Amercaeins ridens
https://www.ephemeroptera-galactica.com/pubs/pub_p/pubprovonshaa1985p1.pdf

Provonchsa & MaCafferty Amercaenis cusabo
https://www.ephemeroptera-galactica.com/pubs/pub_p/pubprovonshaa2006p1.pdf

Funk & Sweeny Eruylophella Larva.
http://www.ephemeroptera-galactica.com/pubs/pub_f/pubfunkd1994p209.pdf

apobaetis/ porcloeoids/ paracloeon/ procloeon/ cloeon,
http://www.ephemeroptera-galactica.com/pubs/pub_d/pubdayw1955p121.pdf

Mcc Pentagena
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1972p51.pdf

Biosystematics of Danella Mcc
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1972p51.pdf

Mcc / Nixe / Leucrocuta, hepta / larva
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1977p350.pdf

Moribaetis mimbresaurus mexico
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw2007p696.pdf

Caenis arwini / Texas
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw2001p349.pdf

Dicrephemera new subgenus of Ephemera
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1973p300.pdf

Baetis mimbresaurus SW NA
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw2007p696.pdf

Moribaetis macaferti Mexico-SA
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1998p117.pdf

NA / Baetis macdunnoughi Ide
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1979p26.pdf

Baetiods in Texas
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1993p59.pdf

Phylogenetics and the classification of the Timpanoga complex
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1994p569.pdf

Systematics and biology of Acentrella turbida
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1994p301.pdf

Western Rhithrogena
http://www.ephemeroptera-galactica.com/pubs/pub_a/puballenr1978p311.pdf

Alemletus revision
http://www.ephemeroptera-galactica.com/pubs/pub_z/pubzlotyj1997p251.pdf

Mcd Ameletus 1936 suffusus
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmcdunnoughj1936p207.pdf

Eurylophella larva Funk and Sweeney
http://www.ephemeroptera-galactica.com/pubs/pub_f/pubfunkd1994p209.pdf

Dr Jeff Webb 2002 Masters thesis text part 1 and 2. in this link there is the appendices and all the figures is the first, the second is the text description. Jeff said yes please share it.

https://www.dropbox.com/sh/uqqulrbjtx2f7qq/AADvB9SPUdMkhA3mUTx9f5z_a?dl=0

Heptageniidae of EU
https://observation.org/taxa/12046/

F P Ide 1937 Baetis manual
http://www.ephemeroptera-galactica.com/pubs/pub_i/pubidef1937p219.pdf

Spieth 1941 Hexagenia manual
http://www.ephemeroptera-galactica.com/pubs/pub_s/pubspiethh1941p233.pdf

Isonychia manual;
http://www.ephemeroptera-galactica.com/pubs/pub_k/pubkondratieffb1984p129.pdf

Berner 1948; Isonychia tusculanensis Berner, 1948 [USA:SE]
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubbernerl1948p117.pdf

Ecdyonurus; spain.
https://www.academia.edu/14781111/The_Status_of_Knowledge_of_the_Genus_Ecdyonurus_in_the_Iberian_Peninsula_with_Description_of_Two_New_Species_of_the_E_venosus_Group_from_Spain_Ephemeroptera_Heptageniidae_?email_work_card=view-paper

Jeff Webb 2008 Heptagenia of the world part II, PhD thesis
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2008p1.pdf?fbclid=IwAR1fhpHHYvotB02ORdJLclMO3T2QVTKjQDjtS1zmHelYJH9RpNJW3BAOgAk

Bednarik & MaCafferty 1979 biosystematics of Stenonema and and now Maccaffertium mayflies;
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubbednarika1979p1.pdf?fbclid=IwAR0e8aCfLLDvGZ2CDC2nZFMcT_R9JtJXLeKevRx9CAnvVIlDtKK_Cu7zPPI

Lewis P A 1974; the taxonomy and ecology of stenonema mayflies;
http://www.ephemeroptera-galactica.com/pubs/pub_l/publewisp1974p1.pdf?fbclid=IwAR2pnWIebMjk2sYYmmtfiEtlwFjx77LixPJ4nuZ55v5FZ7PBqkebkAII0UQ

Fly Fishing Entomology spicies update;
http://www.flyfishingentomology.com/Updated%20Taxonomy%20for%20The%20Biology%20Of%20Mayflies.htm?fbclid=IwAR3TQx9pgsNjh3p4z5ajqzOGdlHfgmJV6EVc7W8w2MUV28LmfC7n2uRmaXw

Afrotropical and Oriental Genus Afrunus
http://www.ephemeroptera-galactica.com/pubs/pub_s/pubschoonbeeh1968p1.pdf

Callibaetis / SW America / nsp C distinctus
http://www.ephemeroptera-galactica.com/pubs/pub_l/publugoc1996p1.pdf

Texas Baetidae / Callibaetis
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1993p59.pdf

Webb 2006; Compsoneuria
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2006p55.pdf

Webb 2003 Heptagenia adaequata
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2003p289.pdf

Webb 2007 Heptagenia Walshi
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2007n63p1.pdf

Webb 2007; Heptageniidae of Borneo
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2007p41.pdf

Webb 2006; Epeorus aculeatus, oriental range
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2006p65.pdf

Webb 2007; NE; Epeorus
http://www.ephemeroptera-galactica.com/pubs/pub_w/pubwebbj2006p57.pdf

Cloeodes of Argentina 2008 Carolina Nieto*
http://www.ephemeroptera-galactica.com/pubs/pub_n/pubnietoc2008p1.pdf

Carolina Nieto* September 2008 Callibaetis larva / adults south America
http://www.ephemeroptera-galactica.com/pubs/pub_n/pubnietoc2008p229.pdf

Callibaetoides of Mexico 2013 Paulo Vilela Cruz
file:///C:/Users/home/Downloads/A_new_genus_and_species_of_Baetidae_Insecta_Epheme.pdf

Ephemeroptera thalerospyrus; Chonlakran Auychinda 1 , Luke M. Jacobus 2 , Michel Sartori 3,4 and Boonsatien Boonsoong July 2020;
https://europepmc.org/backend/ptpmcrender.fcgi?accid=PMC7563720&blobtype=pdf

Heptageniidae of Thailand
https://zookeys.pensoft.net/article/3714/

New Zealand; Leptophlebiidae
https://www.academia.edu/32624640/Leptophlebiidae_Insecta_Ephemeroptera_

Costa Rica Leptphlebiidae / genus Ulmeritoides
https://www.academia.edu/882513/New_species_and_records_of_Ulmeritoides_Ephemeroptera_Lep_tophlebiidae_from_Costa_Rica

American genus Farrodes Leptophlebiidae CA
https://www.academia.edu/29059962/Systematics_cladistics_and_biogeography_of_the_American_genus_Farrodes_Ephemeroptera_Leptophlebiidae_Atalophlebiinae_

European Heptagenia
http://www.ephemeroptera-galactica.com/pubs/pub_s/pubsavolainene1991p201.pdf

Maccaffertium lenati
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmccaffertyw1990p760.pdf

Keys to NA mayfly larva
http://dep.wv.gov/wwe/getinvolved/sos/documents/benthic/british_mayflykey.pdf

Mayflies of florida, Lewis Berner
https://ufdc.ufl.edu/AA00032554/00001/1j

Beaty 2010 mayflies on NC
https://files.nc.gov/ncdeq/Water%20Quality/Environmental%20Sciences/BAU/Benthos%20Reference/BAU%20Taxonomy%20-%20Ephemeroptera%20-%2021Oct13%20Full%20version.pdf

Keys to mayflies of florida larva
http://www.ephemeroptera-galactica.com/pubs/pub_p/pubpescadorm2004p1.pdf

Flowers 1980 Heptageniidae split / Heptagenia / Leucrocuta / Nixe
http://www.ephemeroptera-galactica.com/pubs/pub_f/pubflowersr1980p296.pdf

Funk $ Sweeney 1994 Eurylophella larva
http://www.ephemeroptera-galactica.com/pubs/pub_f/pubfunkd1994p209.pdf

BD Burks 1953 Ephemeroptera of Illinois
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubburksb1953p1.pdf

Burain Epeorus frisoni 2008
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubburians2008p277.pdf

SK Burian 2002 Eurylophella coxalis
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubburians2002p602.pdf

BD Burks 1946 new Heptageniidae
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubburksb1946p607.pdf

McDounngh 1925 Epeorus
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmcdunnoughj1925p168.pdf

2003 Chinese Ephemera
http://www.ephemeroptera-galactica.com/pubs/pub_z/pubzhouc2003p69.pdf

Important Chinese Epeorus melli 2007;
http://www.ephemeroptera-galactica.com/pubs/pub_z/pubzhouc2007p49.pdf

Important Heptageniidae of Korea;
http://www.ephemeroptera-galactica.com/pubs/pub_y/pubyooni1984p1.pdf

McDounngh 1933 nepotellum / mediopunctatum
http://www.ephemeroptera-galactica.com/pubs/pub_m/pubmcdunnoughj1933p16.pdf

F P Ide 1935; post embryonic study of larvae development;
http://www.ephemeroptera-galactica.com/pubs/pub_i/pubidef1935p433.pdf

Anepeorus larva; Beckett 1977.
http://www.ephemeroptera-galactica.com/pubs/pub_b/pubbeckettd1977p240.pdf

Baetidae keys;
http://www.usu.edu/buglab/Content/Western%20Baetidae%20Nymphs.pdf

Stonefly website.
https://theearlybirder.org/stoneflies/?fbclid=IwAR2Nbvn-iLNY6D1bYC_KfsGUKfmfFjR3EI2i2AufcKjeUhSZY9t0rYo-0zQ

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